How can PHP developers optimize database queries to improve performance?

PHP developers can optimize database queries by using indexes on columns frequently used in WHERE clauses, avoiding SELECT * and fetching only necessary columns, using prepared statements to prevent SQL injection attacks, and minimizing the number of queries by using JOINs instead of multiple queries. Additionally, developers can use caching mechanisms like Memcached or Redis to store frequently accessed data and reduce database load.

// Example of optimizing a database query by using an index on a frequently used column
$query = "SELECT * FROM users WHERE username = :username";
$stmt = $pdo->prepare($query);
$stmt->bindParam(':username', $username, PDO::PARAM_STR);
$stmt->execute();